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Actually... I believe I am qualified to answer this question. Here we are discussing first-generation infrared seekers:
The core component of a first-generation infrared guidance system is a partially blackened transparent plastic disk known as a reticle. The image shows one type of reticle, called the "rising sun reticle" (translated from Chinese; I am not sure of its foreign name).
In a missile seeker, the following parts are present:
[Optical components] – [Reticle] – [Single photosensitive element]
You can see that half of the reticle is completely blackened, while the other half has evenly spaced stripes.
Now, a target appears in the field of view. With only a single photosensitive element, we cannot directly extract the target's angular information. The reticle, however, intelligently modulates the continuous signal to tell the guidance system where the target is.
The target signal passes through the optical components and is projected onto the reticle. Through the reticle, the target causes a flickering effect. From this flicker, the guidance system behind the photosensitive element can extract information. The specific method is as follows:
First, we need to break down the angle between the missile and the target. Here, we decompose it into the angle between the missile's axis and the target, and the angle formed on the plane normal to the missile axis relative to a reference direction (the specific direction is irrelevant, as long as it is fixed relative to the missile).
When the light source falls on the striped region, it produces a continuously flickering signal. If the target deviates significantly from the missile's axis, the flickering signal will have a higher amplitude; if the deviation is small, the signal amplitude will be lower.
When the light source falls on the blackened semicircle, the guidance system uses this period of prolonged signal interruption, along with the rotation speed and position of the reticle, to calculate the angle on the missile's normal plane relative to the reference direction (again, the specific direction is irrelevant, as long as it is fixed relative to the missile).
By using the two parts of the reticle to calculate these two angles separately, the complete angular information of the target is obtained.